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One Virtualized Domain Controller Cannot Process Group Policy

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I have 3 DCs, 2 sites.  Main site 1 DC standalone hardware, 1 DC Hyper V guest, remote site 1 DC Hyper V guest.  Hyper V guest DC at main site will not process GP with error 1096 and only error 1001 in the application log. 

DC has all functionality except that it cannot browse it's own shares or any other network shares.  All other computers in the forest can browse the shares on this DC fine, the DC itself just cannot access the SYSVOL share to process GP.  It can surf the internet and ping the world.  It's not DNS.

The DC was working for years without issue.  All of a sudden just lost internet connectivity.  I removed and readded both the NIC and the virtual bus and that resolved the internet issue, but the DC still will not process GP.  I can go to the DC's C: drive and browse to see that the SYSVOL structure is there and intact.

I've checked registry entries and everything else I can think of.  I am reading posts that people have unjoined/rejoined, reinstalled, etc., and the issue persists.  All other Hyper V guests on two hosts across two sites work without issue.  I have no idea why this DC's NIC just stopped being able to browse.  This is not a new environment, nothing new was introduced to cause the problem.

Any assistance would be appreciated.  DCs are Windows 2008 R2.  






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